Year of Publication: 1949, Vol. 12 (02)

Date Published 07 December 1949
Holttum, R.E.
The Selection Type-Species of Some Old Genera of Ferns [Page 303 - 306]
Abstract:
Some early genera of ferns are so heterogeneous that the authors cannot have had clear ideas in founding them. It is sometimes possible to exclude certain species as not conforming  to the author's generic description, but it is often not possible to say that one species rather than another is clearly indicated by that description. The generic description is thus not an infallible guide in the selection of a type-species.  The usual procedure has been to follow the first author who divides a genus. The process may be repeated more than once. Often the authors who split genera did not indicate a type-species; one can only agree that the type-species must be contained in that part of the original genus which is retained under the original generic name. It may occur that subsequent authors in dividing a genus do not pay attention to the original generic description, and so may retain under the original generic name a species that does not well agree with that description, removing all others to another genus or genera. In such cases it may seem reasonable to select a new type-species of the original genus. But if ( as often happens) none of the original species correspond uniquely to the description, there may well be difference of opinion as to which species most nearly corresponds to it. The result will be confusion. I suggest that, in the interests of uniformity of nomenclature, it is better to follow the first divider of a genus, even though he may not select (by implication or otherwise) a type-species that most nearly corresponds to the original description. This may not be ideal arrangement; but in practice no ideal arrangement is possible, and I suggest that it is better to follow a definite rule rather than an indefinite one which will allow of differences of opinion and, as a result, confusion of nomenclature. On the other hand, there should certainly be a recommendation that a later author, in dividing a genus, should principally consider the generic description when selecting a type-species. Linnean genera can be treated like any others, having regard to the convention that the species described in 1753 are associated with the generic descriptions of 1754, and to the rule that legitimate nomenclature begins in 1753 (art. 20).  I apply these principles to the names of certain fern genera as follows:      

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Furtado C. X.
A Further Commentary in the Rules of Nomencalture [Page 311 - 377]
Abstract:
In my previous commentary on the laws of botanical nomenclature (Gard. Bull. Straits Settl. IX, 1937, pp. 223 - 284), an attempt was made to clarify a number of provisions which appeared vague or inconsistent in the 1935 Rules. Later some amendments were proposed by me for consideration at the 1940 Botanical Congress on points that appeared to me the most essential for future progress (Gard. Bull. Straits Settl. XI, 1939, pp. 1 - 30). However, I venture here to isuue a complementary set of proposals and a commentary in the hope that the philosophical basis of nomenclature may receive due consideration in the revision of the Rules at the 1950 Congress. I submit that the time has come when botanists should pause to analyse first principles involved in the system of the rules as a whole and then examine the different rules accordingly. I propose therefore that a special Committee be appointed to consider in detail the principles involved.  Should this Committee agree to a principle, but not to the location or the form of an amendent embodying the principle - two reasons why a good proposal may be rejected - the Commitee should be empowered  to suggest a better place and/or better wording so that the principle might be incorporated in the code.

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Furtado, C.X.
Palmae Malesicae - X, The Malayan Species of Salacca [Page 378 - 403]
Abstract:
Of the 13 species of the genus, the following are wild in the Malay Peninsula : S. affinis, S. conferta, S. flabellata, S. glabrescens, S. Rumphii and S. Scortechinii. RIDLEY does not include the last mentioned species in his Flora (1925), nor does he give any reason for this omission. Beccari's plate of the type of S. Scortechinii looks like a mixture consisting of a young leaf of S. affinis and a spadix of S. conferta, but I do feel justified in making this reduction without being able to compare the original material with recent collection.  Salacca conferta is split into two species by BURRET (1942), under the genus Eleiodoxa, as E. conferta and E. orthoschista, the latter based on material collected in Singapore.  Though we have very good material from Singapore, we have very little from Malacca, the type locality of S. conferta ; on the evidence at present available, I am not able to separate the two species. S. flabellata is the only new species described here; it is reported to be very common in two places in Kemaman (at Sungei Nipah and at Bukit Kajang). It is the smallest species in the genus and is easily recognized by its individed leaves, a character not found in any other Salacca except in seedling stages. The species is known from male specimens only.  In the key to the species, the non-malayan species S. sumatrana, S. vermicularis and S. edulis, are given in order to make clear the identity of S. edulis, which is found occasionally cultivated or as an escape in Malaya, and is probably the species commonly cultivated in Java for the export of fruits.  I have also given the synonyms of S. edulis, so that the use of the name is clarified. S. borneensis has been reduced to a variety of S. affinis, but the variety has not been recorded in Malaya.    

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Year of Publication: 1949, Vol. 12 (01)

Date Published 30 April 1949

Year of Publication: 1947, Vol. 11 (04)

Date Published 30 September 1947
Holttum, R. E.
New Species of Vascular Plants from the Malay Peninsula [Page 267 - 298]
Abstract:
During a survey of the ferns, orchids, grasses and Cyperaceae of the Malay Peninsula, based on all specimens in the Singapore herbarium, a number of undescribed species were found. These are described below. It is hoped that full revised accounts of all these families, as represented in the Malay Peninsula, will be published later. Various changed of names, found necessary during the work, are also here published.

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Henderson, M. R.
New Species of Eugenia Linn. (Myrtaceae) [Page 299 - 338]
Abstract:
The following new species of Eugenia are described in anticipation of a critical review of the genus in the Malay Peninsula which it is hoped will be published in the near future.

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Year of Publication: 1941, Vol. 11 (03)

Date Published 30 August 1941
Furtado, C. X.
Alocasia macrorrhiza and its Varieties [Page 244 - 257]
Abstract:
The binomial Alocasia macrorrhiza  has been used in more than one sense.  Some of the plants which have passed under this name are economically very important and found in cultivation or as escapes in the tropics of both hemispheres; others are of little economic value and not known in cultivation or as escapes except in their native country and perhaps in some botanic gardens. Further, many experiments and analyses have been made in order to test to their value as food to men and animals, and the results have been sometimes conflicting. The existence of many varieties in some of the species called by this name is also a factor that has caused further ambiguity. The present paper is an attempt towards the clarification of this ambiguity. It is my pleasant duty to record here my thanks to Dr. R. C. BAKHUIZEN VAN DEN BRINK, Agricutural Deaprtment, Buitenzorg, Dr. G. J. A. TERRA, Horticultural Division, Batavia, Dr. C. G. G. J. VAN STEENIS, Botanic Gardens and Herbarium, Buitenzorg, Dr. K. BISWAS, Royal Botanic Gardens, Calcutta, and Mr. E. F. ALLEN, Agricultural Department, Teluk Anson, Perak, for procuring for me planting material of the species and varieties connected with the nomenclatural problems of Alocasia macrorrhiza and its varieties ; without this material it would have been impossible for me to clear many difficulties discussed in this paper.

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Year of Publication: 1940, Vol. 11 (02)

Date Published 29 November 1940
Furtado, C. X.
Palmae Malesicae, VIII - The Genus Licuala in the Malay Peninsula [Page 31 - 73]
Abstract:
The results embodied in this paper are of a study undertaken with the view of arranging systematically the Singapore herbarium specimens of the Licuala species indigenous in the Malay Peninsula. The present world conditions, entailing the cessation of intercouse with certain botanical instituitions, the dislocation of the staff at some others, and the risks to parcels during transit across oceans have precluded my obtaining either critical notes of the type specimens preserved outside the Malay Peninsula, or a loan of the specimens themselves. Under such limitations the results of my undertaking would have been of little systematic value but for the valuable assistance derived from BECCARI"S monographic work on the genus published first in an abbreviated form in Webbia V, 1921, pp. 22 - 55, as a part of Recensione delle Palms del Vecchio Mondo appartenenti alla tribu delle Corypheae and then in an extended form in the Annals of the Royal Botanic Gardens, Calcutta, XIII, printed in 1931 and published in 1933 ( cited below as Calcutta Annals or Calc. Annals XIII, 1933 ). In this latter work BECCARI incorporated the results of the opportunities he had not only of consulting the type material of most of the species he studied and of comparing newer material with it, but also of making lengthy critical descriptions, analytical drawings and photographic plates of such specimens.

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Holttum, R. E.
The Fern Genus Diplazium in the Malay Peninsula [Page 74 - 108]
Abstract:
The ferns of the genus Diplazium occurring in the Malay Peninsula are all, with the exception of D. esculentum and D. polypodioides, found on the ground in shady forest, many of them only in moist places near streams. They are especially characteristics of the valleys at medium elevations in the mountains, though a few species are common in the more open parts of the forest. There are no epiphytes and no climbing species among them. Nearly all have short erect rootstocks ( sometimes developing a short trunk) and tufted fronds, sterile and fertile fronds being almost alike. There is comparatively little difference of external form among the species of the same degree of division (i.e., among pinnate and among bipinnate species), and there is ofcourse a certain amount of variation according to age and influences of habitat, so that the clear distinction of species is at first sight difficult, and there has in fact been much confusion in the past. I have found, however, that the characters of the scales are most useful in distinguishing species of similar habit, and I believe that I have arrived at a clear distinction of our local species, with the possible exception of D. Prescottianum. I have been unable to identify four of our species with any described from places outside the Peninsula, and so have described them as new. There are undoubtedly a very large number of species within the Malayan region, but I think that a good many have been described more than once. The descriptions are, however, inadequate as a basis for comparison, and much herbarium material is incomplete (part only of a leaf, or lacking in scales, or representing an immature stage). A study of much material over a wide area is necessary before a proper account of the genus in the Malayan region can be given. Diplazium cordifolium provides a remarkable case of polymorphism leading to confusion of names. Young plants develop rather large simple fronds which are often fertile, and this stage may persist indefinitely, especially in the lowlands. Mature plants, however, often bear pinnate fronds with several pairs of pinnae. The two stages have been separately named ; and the fact that other superficially similar simply pinnate species have sometimes a few veins anastomosing has also led to confusion between these and the mature form of D. cordifolium. The young stages of species may be characteristic, and so useful diagnostically, in this genus as in other fern genera, but they have been little srudied. I have noted especially the early stages of D. bantamense and D. riparium. In the following account of the species found in the Malay Peninsula, I have not attempted to give full lists of synonyms. There has been so much confusion in the application of names that such a lists of synonyms would be difficult to compile accurately. I believe that the names used are all correctly applicable to our species ; but I differ in some cases from the usage of the recent book by Backer and Posthumus on the ferns of Java. In such cases, the synonymy is briefly discussed. As regards citation of specimens, this is done only where I believe there has been confusion in the past, or in the case of new species ; in cases of well characterized species such as D. accedens and D. cordifolium there seems no need for the publication of lists of specimens. Specimens collected in our Singapore Field Number series (started by Burkill in 1912) are cited as S.F.N. Duplicates of many of these will be found in other herbaria, in some cases under other names than those now given. A list of numbers at the end of this paper may therefore prove helpful. It should be noted that the earlier distribution labels did not bear the words "Singapore Field Number". A fairly full description is given for each species. To aid in the identification of specimens, the most distinctive characters in each description are italicised.

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Jagoe, R. B.
Carpet Grass, Axonopus spp. [Page 109 - 118]
Abstract:
For many years the apparent differences between broad-leaved and narrow-leaved forms of Axonopus compressus, or Carpet grass, has not been generally regarded as of specific rank, although the names Paspalum platycaule Poir, for the broad-leaved form and Paspalum compressum, for the narrow-leaved form, have been used by some persons. The original Paspalum compressum, Swartz., is however, undoubtedly the broad-leaved grass. These two Carpet grasses have now been recognised as distinct species, the broad-leaved form retaining the present name ( Axonopus compressus Beauv.), while Miss Agnes Chase, an American authority, has defined the narrow-leaved grass as a new species, and named it Axonopus affinis, as follows :- Axonopus affinis, sp. novo.

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Holttum, R. E.
Periodic Leaf-Change and Flowering of Trees in Singapore [Page 119 - 176]
Abstract:
In my earlier paper on the subject, I published observations made on a number of trees during the years 1927 - 1931, and summarized previous observations published elsewhere on the behaviour of trees in the eastern tropics. I also gave a short account of the climate of Singapore. The main items of new information arising from my own observations was that a number of deciduous trees had fairly constant leaf-periods, which had no relation to the yearly calendar; and the conclusion drawn from this was that the length of leaf-period in such trees is due to primarily to senescence of leaves, the climate of Singapore being so uniform as to have little or no control. Subsequent observations, in some cases over a total period of ten years or more, have confirmed the fact that many deciduous trees have fairly constant leaf-periods not related to climate; but they have also shown many curious irregularities which are not always easy to explain. It is clear that there is every variety of response from almost complete independence of climate change to extreme susceptibility, and a tree that is regular in behaviour for a time may later show an unexpected change (e.g. Koompassia).  My earlier generalizations therefore need modification in some measure, and a fuller discussion of the whole subject is given below, followed by details of behaviour of individual trees.

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