Year of Publication: 1955, Vol. 14 (2)

Date Published 15 February 1955
Sinclair, J.
A Revision of the Malayan Annonaceae [Page 149 - 516]
Abstract:
No major publications on the Malayan Annonaceae have appeared after King and Ridley's accounts. Since their time much herbarium material has accumulated and it is now desirable to revise the family. We should be grateful to these authors and to all the other pioneers for their writings on the subject.  We are now more fortunate than they since this lapse of time has placed at our disposal more facts, more material and new scientific concepts. The prresent is another account but it cannot pretend to be comprehensive. There are still too many disturbing and puzzling questions to be solved but it is hoped to solve some of them in the future. Where more information is required or where certain statements appear to be doubtful, then such points are mentioned in the notes after the description of the species. There are still some imperfectly known species lacking either flower or fruit. A good deal of living material has been seen when such was available, it was possible to add colour notes not mentioned in any of the text-books or papers. More material is still required. There are several species which have only once been collected and it is essential to know more about their distribution. Flowers are more important than fruits for identification while sterile material is sometimes not of much use. In some cases as is pointed out later, it is of no value.  

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Furtado, C.X.
Palmae Malesicae - XVIII, Two New Calamoid Genera in Malaya [Page 517 - 529]
Abstract:
While studying specific affinities in the genus Calamus with the view to revising the Malayan material in the Singapore herbarium, a few species were noticed to be so anomalous as to justify their separation from Calamus; and as they do not form a uniform group, two new genera have been proposed here, namely Cornera and Schizospatha. The first genus is named after Dr. E. J. H. Corner of the University of Cambridge, England, who, when Assistant Director of the Botanic Gardens, Singapore contributed much to our knowledge of the Malayan vegetation in general; the second name has been coined to emphasize the fact that in this calamoid genus the spadix branches emerge by puncturing the spathes. The species in both these genera are climbing and their spadices are much abbreviated, in Cornera terminating each in a short filiform appendix.  None of the species are cirriferous, but one species in Cornera (C. conirostris) produces a short cirrus which however bears also abbreviated leaflets to make the leaves sub-cirriferous in the terminology adopted for describing rattans.  The primary spathes, though in texture and armature reminding one of Daemonorops and of its relatives, are conspicuously tubular at least in part, the leaf-sheaths are flagelliferous and the female calyx is as nearly as long as the corolla - three characters that show the clear affinities of these two genera with Calamus. The primary spathes in Cornera are armed, coriaceous, ventricose or inflated towards the apex which in each spathe terminates often in an ear-shaped limb having a long or short beak. The primary branches of the spadix have each a stout axis which, growing ina straight line with the main axis below, appears as if it were the continuation of the latter, while the main axis above is so much more slender than the branch-axis that it appears to be a true branch - characters not known to ocur in any other calamoid genus.  The axes of the spikelets which are short and congested in each spadix-branch, are also thick; and the flowers (male and female) and the fruit are also very much larger than in any other rattan genus. The fruit scales are not channelled in the middle. The spadices in Schizospatha are much more abbreviated than those in Cornera and do not have a filiform appendix at the end. The primary spathes are papyraceous, often fragile, imbricate, longer than the included internode and the axillary branch, and gradually shorter towards the end of the spadix; this means that each spathe covers partly or entirely the one above, and that the longest internode and the largest spathe are the lowermost in the spadix, and the shortest are at the apex.  Often the terminal portion of the spadix is abnormal; this may contain two or more spathes which though amplexicaul at the base, are open and cymbiform, each subtending in its axil an abortive or fertile spikelet. This entire abnormal part is wholly enclosed, before anthesis, in a large cymbiform, amplexicaul spathe. The tubular primary spathes do not dehisce but remain closed eo that the spadis branches with their spikelets emerge by puncturing their respective axillant spathe on its dorsal side, a mode of orientation for sssspadix branches not known in any other rattan genus. Later, as the spikelets develop and the spadix bends, the spathes become torn and appear to have dehisced naturally, but the basal spathes will often reveal the true mode of emergence of the spadix branches. The spathes maybe entirely unarmed or occasionally the lowermost spathe is armed at the base and obscurely so in the lamina.          

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Burkill,  I. H.
William Farquhar's Second Book of Drawings of Malacca Plants [Page 530 - 533]
Abstract:
A short account was given in 1949 (this bulletin 12, pp. 404 -407) of a book of illustrations of Malacca medicinal plants presented in 1827 to the Royal Asiatic Society by Lieut.-Col. William Farquhar.  A second book exists and is the subject of this note.  William Farquhar, it will be remembered, was the first resident and Commandant of Singapore; he had been resident of Malacca previously and while there had employed a Chinese artist to make for him illustrations of useful plants in an attempt to learn to know them. His second book may be divided into three sections; the first rattans, the second results of an ascent of Mount Ophir, and the third trees valuable for their timber or resins. I have attempted to identify his plants portrayed as there is an interest in knowing on what jungle produce Malacca was living. The artist excelled in painting foliage; and twigs must have been brought to him for the purpose. Some of the trees he may have known in the forest but he did not attempt to paint them from life; instead he drew and coloured trunks and branches in what maybe called diagrams. There is only one representation of a flower and that erroneous and three of the fruit. A Malay wrote in arabic characters the plant names except where I indicate this below. These names and the foliage are in truth all that a botanist has to guide him in determinating the plants.  Farquhar was proud of his attempt and took the drawings with him in December 1818 when he went under instructions to join Raffles at Penang; and William Jack was shown them, who commented to Wallich in a letter that 'they are deficient in many essential points .... but will be extremely useful as a guide, by taking the native names .... and making enquiries accordingly'. Farquhar later showed them to Wallich who made some shots at naming a few. This would be in 1822 in which year Wallich resided for a short time in Singapore. Someone, propably Farquhar himself, showed them to Lindley whose handwriting is against one. This would be in London and just before thery were given to the Asiatic Society.

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Garrard, A.
The Germination and Longevity of Seeds in an Equatorial Climate [Page 534 - 545]
Abstract:
Singapore is situated 1 17" North latitude on the South side of Singapore Island. It has very uniform temperature, high humidity and copious rainfall (Colony of Singapore Annual Report). The mean temperature of the coolest month (December) is only 3F lower than the hottest (May). The absolute minimum is 70F and the absolute maximum 93F, but these are rarely reached. The normal range is 75 - 80F on a wet day and 74 - 89F on a dry day.  (Holttum, 1953). The rainfall is between 85 and 118 inches per year. Its distribution varies from year to year, but there is a maximum fall in the months December - January and generally a short period of drier and windy weather in late January or February. However, a month in which less than 2.5" rainfalls is rare, and occurs about every two years in the February - March or July - September periods.  Crocker and other workers at the Boyce Thompson Institute have made extensive investigations on the conditions most favourable for the storage of seeds and have found that most seeds require a low temperature and a low moisture content. (Crocker, 1948). The climate of Singapore maybe expected to be unfavourable since the temperature and humidity are relatively high.  Moreover, Singapore is characterised by having a continuous growing season. During the whole year the conditions are favourable for germination. The ability for seeds to undergo a period of dormancy during unfavourable conditions (as in temperate or monsoon climates) is not an essential feature for the survival of plants grown here. However, a large number of locally grown plants have been imported from other more seasonal climates and have seeds which normally undergo a resting period before germination takes place. Some such seeds may require a period of 'after-ripening' before germination can occur. It may be that the climate is not suitable for this to take place and seed propagation of such plants cannot occur here.  It is propable that the majority of seeds germinate shortly after they reach the ground, but it is interesting to investigate the survival period of those which may have fallen in a place unfavourable for germination. Such results are also interesting from the horticultural point of view since a gardener may not find it convenient to plant freshly gathered seed immediately. Seeds which do not degenerate under Singapore conditions in storage can be safely stored until required.      

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Year of Publication: 1953, Vol. 14 (1)

Date Published 01 August 1953
Sinclair, J.
Additions to the Flora of Singapore I [Page 30 - 39]
Abstract:
The following paper gives a list of thirty-seven species new to the flora of Singapore. Twelve of these are new to the flora of Malaya as well.  Seventeen are native and twenty are not native. Of the twelve new to Malaya, three are native and nine are not. Some of the species new to Singapore were not actually obtained in the island itself but in the small islands situated to the south which are included in the Colony of Singapore. The nearest island to the coast of Singapore is Pulau Samulun, roughly about one-eighth of a mile away. The second nearest is Pulau Damar Laut, about a quarter of a mile distant. The farthest, Pulau Pawai and P. Senang are some eight and nine miles away. Of the thirty seven species, eleven were found in these islands and are not known to occur in Singapore. A further four were found in these islands but they also occur in Singapore. Of these fifteen are native and five not. The majority of these islands do not appear to have been visited by botanists since there are no plant records from them mentioned in Ridley's Flora and further there are no herbarium specimens preserved in the Herbarium of the Singapore Botanic Gardens.  This is probably the reason why they have yielded such a large number of new records.  One of them, Pulau Busing  where Cordia subcordata was obtained, is a mere strip of uninhabited mangrove which one might think scarcely worth while visiting. In the flora of these islands we find some of the elements of the east coast flora of Malaya and it is possible that seeds of some of the species were carried by the sea from the north. This paper also deals with a few species which were collected long ago and which are now probably extinct in their former localities. New localities are given for them. Finally there are four species which have ben previously recorded but there are either no exact localities or no herbarium specimens preserved.      

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Sinclair, J.
Notes on Siamese Annonaceae [Page 40 - 44]
Abstract:
In the course of a revision of Malayan Annonaceae, I have at the same time for comparative purpose examined some Siamese Annonaceae kindly lent to me by the Department of Agriculture, Bangkok. As a result there are several new records of species which are not listed in Craib's 'Florae Siamensis Enumeratio'. Several new combinations and nomenclatural changes also have to be made. The species marked with an asterisk are new to Siam.

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Sinclair, J.
Notes on Indian and Burmese Annonaceae [Page 45 - 48]
Abstract:
In the course of a revision of Malayan Annonaceae, I have at the same time for comparative purposes examined a good many Indian and Burmese species. As a result of this certain new combinations are necessary to bring the nomenclature up-to-date. These new combinations as well as some miscellaneous comments now follow.

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Furtado, C.X.
The Genus Daemonorops in Malaya [Page 49 - 148]
Abstract:
Daemonorops is a genus of rotans so closely allied to Calamus that it was often regarded as a section of the latter. This is because Calamus is comparatively a polymorphic genus with sections having characters very near those of Daemonorops. However the one character by which a Daemonorops species maybe readily distinguished from that of Calamus lies in the function of the spadices, which in Daemonorops never serve as climbing organs. Consequently the spadices are generally short, never show a tendency to become long and whip-like or to produce a whip-like appendage (flagellum) at the apex, nor bear on spathes or axes or both, reflexed hooks which are the kind of spines which aid a plant to climb. The spathes moreover all fall off excepting the outermost one which in some cases persists for a long time; even in these cases the persistent basal spathe splits throughout its entire length on the ventral side.  In Calamus, on the other hand, even when the spadices are short and bear no whip-like appendage, the spathes and the axis of the spadix are armed with hooked claws; the spathes are persistent and tubular at least at base. This last character is especially useful in distinguishing from Daemonorops the small species of Calamus which, being acaulescent, may not show even the vestigial hooks and appendices in spadices. In very rare cases, as in C. hypoleucus of India, this tubular base may not be conspicuous because of the shortness of the basal tube and drying out of the spathe; in such instances the fact that the seed albumen is homogeneous also that the secondary spathes are tubular, will clinch the species as of Calamus.  

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Year of Publication: 1950, Vol. 13 (01)

Date Published 01 September 1951
Holttum, R.E.
A New Malayan Vanilla [Page 251 - 253]
Abstract:
Vanilla plants of all species (with the exception of V. aphylla) are vegetatively much alike, and in the south of Malaya they rarely flower. The flowers are also short-lived. The few lowland plants actually seen in flower by botanists have all belonged to the species Vanilla Griffithii, and it has been asumed that this is the only lowland species of Vanilla in this country. Recently however Mr. J. A. le Doux found a flowering  plant near Kota Tinggi, Johore, and this proved to be quite distinct from V. griffithii in the form of the lip.  It is also different from all recorded species from Sumatra, Borneo and Siam. In the structure of the flower, this new species seems to be nearest to V. aphylla, which occurs in the north of Malaya. Flowers of all species of Vanilla have a mass of appendages of some sort in the middle of the lip, opposite the anther. Pollination is affected by small bees which enter the tube formed by the joined lip and column. In returning from the botom of the tube to its mouth, the insect must surmount the obstacle formed by these appendages, and in so doing comes into contact first with the stigma and then the anther. In V. griffithii the appendages consist of very fine twisted wooly hairs. In V. montana Ridl., and V. kinabaluensis Carr, the appendages are a series of thin plates. In the present new species there is a tuft of fine curved but not twisted hairs, directed backwards towards the base of the column. The midlobe of the lip also is covered with thicker erect hairs, as in V. aphylla. The column is joined to the lip for almost the whole of its length, and then the side-lobes enfold the end of the column, which is not visible unless they are folded back.  This arrangement contrasts with that of V, griffithii, in which the column is joined to the lip for only one third of its length, and the side-lobes are reflexed, exposing the end of the column.

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Holttum, R.E.
The Marantaceae of Malaya [Page 254 - 296]
Abstract:
Plants rhizomatous, the rhizome a sympodium, each new element ending in an erect leafy shoot ; rhizome elements usually short. Branches of all kinds, both vegetative and on the inflorescence, bearing first a 2-keeled prophyll backing on to the axis which bears the branch. Erect shoots bearing one to several distichous leaves, the leaves sometimes all basal, sometimes separated by short or long internodes, and usually a terminal inflorescence; in a few cases the inflorescence borne on a separate shoot having only short sheaths, without foliage leaves. Leaf-blade usually elliptic to ovate, nearly always glabrous except sometimes for hairs on either side of the midrib beneath, sometimes with the upper surface variegated, the lower surface sometimes purple; lateral veins oblique, close, fine, with little distinction between main and subsidiary veins; petiole short or long, the portion immediately belwo the blade being thickened and round in section, often slightly curved; sheath short or long, sometimes hairy, the edges usually converging upwards and meeting in a point at the base of the petiole, often without a distinct ligule. Inflorescence always with condensed cymose partial inflorescences in the axils of primary bracts, each successive branch of the cyme enclosed in a 2-3 keeled prophyll, often also with an unkeeled mesophyll opposite the prophyll and closing the gap between its edges. Primary bracts arranged in a simple spike, either distichous or spirally, or with lateral spikes in the axils of lower bracts ; spikes of second and third order sometimes developed. Flowers always paired, the two flowers of a pair usually opening together, but sometimes not, and sometimes unequally stalked, one flower being the mirror image of the other. Ovary inferior, one unilocular or trilocular (trilocular in Malayan species), one ovule in each loculus. Sepals free to the base, usually equal, usually narrow, sometimes persistent on the apex of the fruit. Corolla froming a tube with 3 lobes; lobes usually narrowly triangular or oblong. Staminodes and stamen attached to the corolla-tube, sometimes forming a tube which extends some distance beyond the attachment of the corolla-lobes. Staminodes of the outer whorl two, rarely one; when two, placed on either side of the stamen, petaloid, often unequal, large or small. Staminodes of the inner whorl, two, unequal, called the fleshy staminode and the hooded staminode. Fleshy staminode, small or large, petaloid, usually broad, of thicker texture than the outer staminodes, bearing an oblique fleshy callus on which the stigma rests after release from the hooded staminode. Hooded staminode usually small, with hooded apex and usually a downward-pointing lobe on one side, enclosing the style and stigma until disturbed, and then releasing them. Stamen about as long as hooded staminode, bearing one half of an anther on one side , the other side more or less developed inot a petaloid lamina (Usually narrow). Style and stigma held erect at first by the hooded staminode, when released the upper part springing downwards to form an inverted U, the stigma resting on the callus of the fleshy staminode. Fruit dehiscent or indehiscent, containing 1 to 3 seeds ; in dehiscent fruits the seeds bearing a bilobed basal aril, in indehiscent fruits the aril lacking.  Seed containing a curved or sometimes straight embryo embedded in perisperm, with an opening for germination of the root closed by a special plug as in Musaceae and Zingiberaceae; a hollow ( the perisperm canal ) extending from near the plug into the interior of the seed, in the bay formed by the curve of the embryo, or parallel to the embryo when the latter is straight (Phrynium capitatum).        

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Furtado, C.X.
Palmae Malesicae, XI - The Malayan Species of Korthalsia [Page 300 - 324]
Abstract:
Korthalsias form a small group of clump-forming rattans, the species of which are inadequately known. This inadequacy of our systematic knowledge is due primarily to the inadequacy of herbarium specimens obtained by collectors not well conversant with the needs of the systematist. The one thing that the collector should notice in Korthalsias is the great deal of variation not only in the different parts of an individual stem, but often also in different individual stems of the same clump at different stages. Thus the ocrea of the first leaves on a new stem is somewhat different in texture and in dimensions, and sometimes also in armature, from its condition on the later leaves; after the first few leaves, it soon acquires its definitive form, which is retained by all leaves on the stem except the reduced leaves which are associated with the terminal inflorescence. Similarly in some species the first leaves on a stem are flabellate (undivided, fan-shaped), and are sometimes of a peculiar colour and indumentum (special covering on the surface).  On later leaves the leaflets begin to separate but the terminal two leaflets may remain connate or united. Subsequent leaves may show a gradation of change in shape, size, indumentum, and clour of leaflets and of their stalk-like bases (ansae), and the two terminal leaflets maybe separate; in still later stages the leaves have a clawed whip-like end (eirrus). As the stem approaches its maximal growth and terminates in an inflorescence, the changes in the leaves continue. The main branches of the inflorescence are produced in the axils of leaves showing various gradations in reduction, so that some of the terminal leaves consist each of a leaf-sheath surmounted by a small cirrus, with no leaflets.  Side by side with these changes, there will be noticed variations in the thickness of the stem, the size of the petiole, and in the armature on sheaths, ocrea, etc. However, since the ocrea early acquires a definitive form and changes but very little except in the terminal leaves, it has been found that Malayan Korthalsias can be separated from other rattans both generically and specifically on the adult leaves having ocrea. No other rattan genus has leaves which produce both leaflets which are rhomboidal in shape or have premorse lips (as if bitten off), and long ring-like ocrea at the junction of sheath and petiole. Since leaf-sheaths and petioles are also essential for the correct specific identification of most rattans collectors should therefore be instructed to include a sufficient number of representative bits of stems to show the petioles, sheaths and ocrea.  In order to clarify accurately the status of many species based on juvenile material only, or on material without leaf-sheath and ocrea, it would be useful if sets carefully numbered in the field were made to show all the important variations noticeable in a clump. This should not be taken to mean that the flowering parts are unimportant for a systematic study of Korthalsia; they are useful especially to give an additional confirmation that the identification made on the vegetative characters are correct and also to show their affinities; but it is extremely difficult to identify species on specimens having no ocrea and petioles. Of the inflorescence, spikes and fruits are of very great importance in separating species, but these parts are rarely represented in most specimens.

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Furtado, C.X.
Palmae Malesicae, XII - The Malayan Species of Plectocomiopsis [Page 325 - 338]
Abstract:
Plectocomiopsis is a genus consisting of a group of tufted, climbing palms characterised by the following: The inflorescence is a terminal panicle (monocarpic plants) with its primary branches issuing out of the leaf-axils through the mouth of the tubular sheaths of reduced leaves; the male and female flowers occur in different plants (dioecious); all the spathes including the secondary ones are tubular; the fruits have small scales which are arranged in distinct vertival rows; the seed globose compressed at apex and at base, so as to make it look almost cyclindrical; albumen is homogeneous; and the embryo is basilar. The female flowers are borne either directly on secondary branches, or, like the male flowers, on a small tertiary axis which either remains almost included within its axillant spathel or is developed up to 1 - 3 cm. long.

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Furtado, C.X.
Palmae Malesicae XIII - The Genus Myrialepis [Page 339 - 344]
Abstract:
The name Myrialepis was coined by Beccari to show that the fruits of this genus are distinguished in having innumerable minute, irregularly set scales, a character sufficient to distinguish this genus at once from all other rattans. Its affinities are very close to Plectocomiopsis whose fruit scales are comparatively also very small for rattans, but are much larger than those in Myrialepis, and are arranged in regular rows.  The Myrialepis plants, like those of Korthalsia and Plectocomiopsis, are scandent and tufted, each individual stem terminating its growth with an inflorescence (monocarpic). The inflorescence is either male or female, the plants being dioecious. The side branches of the inflorescence are produced each in the axils of reduced leaves, and emerge each by the mouth of the axillant leaf-sheath ; these branches are divided again once in female plants and twice in male, before they produce spikelets. Leaves are large having no trace of an ocrea at the transition of the sheath into the petiole; they bear a cirrus at the end except when they are produced in the early stages of the stem. Leaflets are linear-lanceolate, provided with a single midrib and many sub-primary ones on both side of the midrib, punctulate with minute scales in the lower surface. Female flowers are solitary at each spathel, with no male or neuter flower attending; each flower has a 3-lobed calyx, a longer 3-parted corolla, a membranous 6-lobed staminodal cup adnate at base to the corolla, and a 3-locular globose ovary. The fruit is one-seeded, globose, and covered with innumerable minute, irregularly set scales.  The seed is slightly broader than long, having a homogenous albumen and a basal embryo ; this last is seated exactly opposite to the chalazal cavity which is punctiform in the cross section of the fruit, and short linear in length. Male inflorescences are more divided than the female, but almost similar; male flowers not seen. The genus is monotypic and was based on specimens collected by Scortechini in Perak. The syntypes which have been reproduced by Beccari (1918) by photographic plates, are from Beccari's herbarium in Florence (Scortechini 513b and sn.).  Hooker f.  (Fl. Brit. Ind. VI, 1893 p. 480 ) who had no access to Beccari's herbarium, quoted Scortechini 457b in Kew Herbarium and failed to make any mention of the actual syntypes on which Beccari had based the genus and the species. A specimen bearing this Scortechini's collection number (preserved in Beccari's herbarium) has been tentatively identified by Beccari (1918) as being a sterile stage of Plectocomiopsis Wrayi. Apparently the genus is widely distributed in Malaya and was at one time very common in Singapore.  Beccari (1918) referred tentatively to this genus some sterile specimens collected from Sumatra, and stated that Plectocomiopsis paradoxus from Burma and P. floribundus from Indon-China might prove to be the species of Myrialepis ; of these two species, however, female flowers and fruit are not known.  

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Furtado, C.X.
Palmae Malesicae XIV - The Species of Plectocomia in Malaya [Page 345 - 350]
Abstract:
The genus Plectocomia, like Korthalsia, Plectocomiopsis and Myrialepis, comprises a group of climbing palms which end their vegetative growth by producing flowers and fruits borne in terminal panicle. In the flowering-stages, however, Plectocomia species are easily separated generically by their large, non-tubular, cymbiform, spathels which cover the spikelets; in young stages of the inflorescence, these spathels are conspicuously imbricate and curl round the axis so as to make the spikes resemble tail-like structures. The side branches of the whole panicle are in the axils of somewhat reduced leaves, and each branch may bear several secondary branches. The spikelets which are produced in the axils of spathels, represent tertiary branches and are shorter than the respective spathels. Male and female flowers are always produced on separate plants; while the flowers in the female are solitary at each notch or pedicel on the spikelet, the flowers in the male plant are always in pairs. The fruit scales are comparatively small and in some species including the one that occurs in Malaya, these scales have a free soft end which makes the fruit squarrorse. The seed albumen is homogeneous and the embryo is basal.  Sterile plants might be easily confused with those of Plectomiopsis and Myrialepis, since the leaves in all these genera have a petiole which is neither gibbose, nor flagellate, and which do not usually produce a conspicuous ocrea (Ocrea is present in P. Corneri) ; the leaflets too are somewhat similar and have thickened margins. However, the small scales which are present in the lower surface of the leaflets in the other two genera, are absent in the leaflets of Plectocomia. Further the stems which are apparently tufted in all the species of Myrialepis and Plectocomiopsis, appear to be solitary in Plectocomia; sometimes the stems in the last mentioned genus produce bulbils or branches near the base, but these apparently do not grow so as to make the plant tufted. Further observations are however necessary to see whether this phenomenon which has been observed on plants growing in the Botanic Gardens, Singaporee, is true also in the wild state. Herbarium specimens of this genus are not well represented in many herbaria, and so collectors should be warned to collect as complete material as possible, together with the notes about the habit of the plant in the field.    

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Furtado, C.X.
Palmae Malesicae XV - The Genus Ceratolobus in Malaya [Page 351 - 359]
Abstract:
Ceratolobus is a peculiar genus among the Indo-Malay-asian palms which have scaly fruits (Lepidocaryeae) and rotan-like stems. As in Calamus and Daemonorops, the spikes are not cycindrical, and the stems do not stop their vegetative growth to produce flowers and fruits and then die, but continue to produce the reproductive parts without ceasing their vegetative growth ; that is, the stem is always polycarpic in this genus. But the character that distinguishes a Ceratolobus species readily from other rotans generically is the peculiar structureof the spadices ; for apart from its being very short and porrect, each panicle of spadix is enclosed in only one external, flattened, lanceolate-fusiform, usually unarmed spathe.  

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Year of Publication: 1950, Vol. 13 (01)

Date Published 30 June 1950
Holttum, R.E.
The Zingiberaceae of the Malay Peninsula [Page 1 - 250]
Abstract:
The only recent critical morphological studies of plants in this family are those by Valeton. Unfortunately, they cover only a limited part of the whole field, but their throughness and clarity of presentation showed the way for further progress. The present work was begun by an examination of living plants of those genera studied by Valeton, and his work is therefore the basis of that now presented. In addition to living plants of species of almost all genera, alcohol material of many other species was available for study, mostly collected during the years 1930 - 1940 by Mr. E. J. H. Corner, with copius field notes, the substance of which is included in the present descriptions. In addition, there are careful coloured drawings of several species described by Mr. Ridley, which have supplied some information not otherwise available. Of some species, however, only dried specimens have been examined, and there are accordingly gaps in necessary information about them. The species descriptions here presented are rather lengthy, but I believe that this is necessary in the present inadequate state of our knowledge of the family. It has been my experience that earlier descriptions frequently omitted data which appeared to me necessary for a proper characterization of the species. For the purpose merely of identifying the species now known to exist in Malaya, much briefer descriptions would be adequate ; but they would not be helpful in the understanding of species still to be discovered, either in this country or in neighbouring territories. I have attempted also a comparative account of the morphology of the inflorescence, which appears to me of basic importance. The present work is confined to species in the Malay Peninsula for two reasons. First, it was mainly prepared in the year 1944, when I had adequate material only of such species for study; and second, I have now other work on hand which prevents me attempting a study of the family over a wider area. Though the work is therefore of necessity a partial one, and though in consequence I cannot offer a satisfactory solution of such problems as that of the typification of Alpinia, I hope that the present work will be a useful basis for that wider study of this interesting family in Malaysia as a whole which is so desirable.  

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Year of Publication: 1949, Vol. 12 (02)

Date Published 07 December 1949
Holttum, R.E.
The Selection Type-Species of Some Old Genera of Ferns [Page 303 - 306]
Abstract:
Some early genera of ferns are so heterogeneous that the authors cannot have had clear ideas in founding them. It is sometimes possible to exclude certain species as not conforming  to the author's generic description, but it is often not possible to say that one species rather than another is clearly indicated by that description. The generic description is thus not an infallible guide in the selection of a type-species.  The usual procedure has been to follow the first author who divides a genus. The process may be repeated more than once. Often the authors who split genera did not indicate a type-species; one can only agree that the type-species must be contained in that part of the original genus which is retained under the original generic name. It may occur that subsequent authors in dividing a genus do not pay attention to the original generic description, and so may retain under the original generic name a species that does not well agree with that description, removing all others to another genus or genera. In such cases it may seem reasonable to select a new type-species of the original genus. But if ( as often happens) none of the original species correspond uniquely to the description, there may well be difference of opinion as to which species most nearly corresponds to it. The result will be confusion. I suggest that, in the interests of uniformity of nomenclature, it is better to follow the first divider of a genus, even though he may not select (by implication or otherwise) a type-species that most nearly corresponds to the original description. This may not be ideal arrangement; but in practice no ideal arrangement is possible, and I suggest that it is better to follow a definite rule rather than an indefinite one which will allow of differences of opinion and, as a result, confusion of nomenclature. On the other hand, there should certainly be a recommendation that a later author, in dividing a genus, should principally consider the generic description when selecting a type-species. Linnean genera can be treated like any others, having regard to the convention that the species described in 1753 are associated with the generic descriptions of 1754, and to the rule that legitimate nomenclature begins in 1753 (art. 20).  I apply these principles to the names of certain fern genera as follows:      

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Furtado C. X.
A Further Commentary in the Rules of Nomencalture [Page 311 - 377]
Abstract:
In my previous commentary on the laws of botanical nomenclature (Gard. Bull. Straits Settl. IX, 1937, pp. 223 - 284), an attempt was made to clarify a number of provisions which appeared vague or inconsistent in the 1935 Rules. Later some amendments were proposed by me for consideration at the 1940 Botanical Congress on points that appeared to me the most essential for future progress (Gard. Bull. Straits Settl. XI, 1939, pp. 1 - 30). However, I venture here to isuue a complementary set of proposals and a commentary in the hope that the philosophical basis of nomenclature may receive due consideration in the revision of the Rules at the 1950 Congress. I submit that the time has come when botanists should pause to analyse first principles involved in the system of the rules as a whole and then examine the different rules accordingly. I propose therefore that a special Committee be appointed to consider in detail the principles involved.  Should this Committee agree to a principle, but not to the location or the form of an amendent embodying the principle - two reasons why a good proposal may be rejected - the Commitee should be empowered  to suggest a better place and/or better wording so that the principle might be incorporated in the code.

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Furtado, C.X.
Palmae Malesicae - X, The Malayan Species of Salacca [Page 378 - 403]
Abstract:
Of the 13 species of the genus, the following are wild in the Malay Peninsula : S. affinis, S. conferta, S. flabellata, S. glabrescens, S. Rumphii and S. Scortechinii. RIDLEY does not include the last mentioned species in his Flora (1925), nor does he give any reason for this omission. Beccari's plate of the type of S. Scortechinii looks like a mixture consisting of a young leaf of S. affinis and a spadix of S. conferta, but I do feel justified in making this reduction without being able to compare the original material with recent collection.  Salacca conferta is split into two species by BURRET (1942), under the genus Eleiodoxa, as E. conferta and E. orthoschista, the latter based on material collected in Singapore.  Though we have very good material from Singapore, we have very little from Malacca, the type locality of S. conferta ; on the evidence at present available, I am not able to separate the two species. S. flabellata is the only new species described here; it is reported to be very common in two places in Kemaman (at Sungei Nipah and at Bukit Kajang). It is the smallest species in the genus and is easily recognized by its individed leaves, a character not found in any other Salacca except in seedling stages. The species is known from male specimens only.  In the key to the species, the non-malayan species S. sumatrana, S. vermicularis and S. edulis, are given in order to make clear the identity of S. edulis, which is found occasionally cultivated or as an escape in Malaya, and is probably the species commonly cultivated in Java for the export of fruits.  I have also given the synonyms of S. edulis, so that the use of the name is clarified. S. borneensis has been reduced to a variety of S. affinis, but the variety has not been recorded in Malaya.    

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